Success!
The Erase, then copy worked. I didn't do the Disk Utility step since this drive had been formatted correctly before the original backup. Option-boot now let me choose the backup as boot disk and it worked normally.
I thought it was interesting that after booting to the backup, the Internal Apple SSD that I had backed up from didn't show up in System Preferences > Startup Disk. The internal drive still worked when I Option-booted back to it.
Thanks for your patient help.
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